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Dr. Steven J. Hageman
hagemansj@appstate.edu
Ph: 1+(828)
262-6609
Fax:1+(828) 262-6503
Department
of
Geology
Appalachian State Univ.
PO. Box 32008
Boone, NC 28608
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Refereed papers, first author:
24. Hageman, S.J.,
Wyse Jackson, P.N., Abernethy, A.,
Steinthorsdottir, M. 2011. Calendar scale,
environmental variation preserved in the skeletal
phenotype of a fossil bryozoan (Rhombopora blakei
n. sp.), Early Carboniferous. Journal of
Paleontology, 85:853-870. 
23. Hageman, S.J., McKinney, F.K., and Jaklin,
A., in press. Testing microhabitat
complexity as a control over bryozoan colonial
growth form and species distributions. Proceedings
of the 15th International Conference of the
Bryozoology Association, Kiel, Germany. 
22. Hageman, S.J., and McKinney, F.K. 2010.
Discrimination of fenestrate bryozoan genera in
morphospace. Palaeontologia Electronica, v. 13 no.
2. 
21. Hageman, S.J., Needham, L.L. and Todd, C.D.
2009. Threshold effects of food concentration on the
skeletal morphology of the marine bryozoan Electra pilosa
(Linnaeus, 1767). Lethaia, 42:438-451. 
20. Hageman, S.J. and Sawyer, J.A. 2006. Phenotypic
variation in the bryozoan Leioclema punctatum (Hall, 1858)
from Mississippian ephemeral host microcommunities.
Journal of Paleontology, 80:1047-1057. 
19. Hageman, S.J., Lukasik, J., McGowran, B. and
Bone, Y. 2003. Paleoenvironmental significance of Celleporaria
(Bryozoa) from modern and Tertiary cool-water
carbonates of southern Australia. Palaios,
18:510-527. 
18. Hageman, S.J. 2003. Complexity generated by
iteration of hierarchical modules in Bryozoa.
Integrative and Comparative Biology, 3:87-98. 
17. Hageman, S.J., Bayer, M.M., and Todd, C.D. 2002.
Partitioning phenotypic variation: implications for
morphometric analyses (Bryozoa). pp. 131-140, In
Wyse Jackson, P., Butler, C., and Spencer Jones, M.
(eds.), Bryozoan Studies 2001. Swets &
Zeitlinger, Lisse, The Netherlands. 
16. Hageman, S.J. 2001. Encrusting invertebrate
faunas and shipwreck histories: ballast stones from
North Carolina Shipwreck 03BUI (Queen Anne's
Revenge). Southeastern Geology, 4:65-73. 
15. Hageman, S.J., James, N.P. and Bone, Y.
2000. Cool-water carbonate production from
epizoic bryozoans on ephemeral substrates.
Palaios, 15:33-48. 
14. Hageman, S.J., Bayer, M. and Todd, C. 1999.
Partitioning phenotypic variation: genotypic,
environmental and residual components from bryozoan
skeletal morphology. Journal of Natural History,
33:1713-1735. 
13. Hageman, S.J., Bock, P.E., Bone, Y. and
McGowran, B. 1998. Bryozoan growth habits:
classification and analysis. Journal of
Paleontology, 72:418-436. 
12. Hageman S.J., Bone Y., McGowran B. and James,
N.P. 1997. Bryozoan colonial growth forms as
paleoenvironmental indicators: evaluation of
methodology and predictive utility. Palaios,
12:405-419. 
11. Hageman, S.J., Bone Y., McGowran B. and James,
N.P. 1996. Bryozoan species distribution on the
cool-water Lacepede Shelf, southern Australia, In
Gordon, D., Smith, A. M., Mackey, J. G., eds.,
Bryozoans in Space and Time: Wellington, National
Institute of Water and Atmospheric Research Ltd., p.
109-116. 
10. Hageman, S.J., Bone Y., McGowran B. and James,
N.P. 1995. Modern bryozoan assemblages
and distribution on the Lacepede Shelf, southern
Australian margin. Australian Journal of Earth
Sciences, 42:571-580. 
9. Hageman, S.J. 1995. Observed
phenotypic variation in a Paleozoic bryozoan:
implications for microevolutionary and systematic
studies. Paleobiology, 21:314-328. 
8. Hageman, S.J. 1994. Microevolutionary
implications of clinal variation in the Paleozoic
bryozoan Streblotrypa.
Lethaia, 27:209-222. 
7. Hageman, S.J. 1993. Effects of
nonnormality on studies of morphological variation
of a rhabdomesine bryozoan, Streblotrypa (Streblascopora)
prisca
(Gabb and Horn). Paleontological Contributions of
the University of Kansas Number 4:1-13. 
6. Hageman, S.J. 1992. Alternative
methods for dealing with non-normality and
heteroscedasticity in paleontologic data. Journal of
Paleontology, 66:857-86

5. Hageman, S.J. 1991. Worthenopora:
a
Mississippian cryptostome (Bryozoa) that looks like
a cheilostome. Journal of Paleontology, 65:648 -661.

4. Hageman, S.J. 1991. Discrete
morphotaxa from a Mississippian fenestrate faunule:
presence and implications. In Bigey, F.P.
(ed.) Bryozooaries actuel et fossile:
Bryozoans living and fossil. Bulletin de la Societe
des Sciences Naturelles de l'Ouest de la France,
Memoire HS 1, p. 147-150. 
3. Hageman, S.J. 1991. Approaches to
systematic and evolutionary studies of perplexing
groups: an example using fenestrate Bryozoa.
Journal of Paleontology, 65:630-647. 
2. Hageman, S.J. 1989. Use of
alumenontos to introduce general paleontologic and
biostratigraphic principles. Journal of Geological
Education, 37:110-113. 
reprinted by
invitation. 1994. Using alumenontos to introduce
evolution and phylogeny. In McComas, W.F.
(ed.) Investigating Evolutionary Biology in
the Laboratory. National Association of Biology
Teachers, Reston, Virginia, p. 111-116.
1. Hageman, S.J. 1988. Image processing
and analysis with microcomputers. Journal of
Paleontology, 62:474-477. 
Refereed
papers, junior author:
5. McKinney, F.K., Hageman, S.J., and Jaklin,
A. 2007. Crossing the ecological divide:
Paleozoic to Modern marine ecosystems in the
Adriatic Sea, The Sedimentary Record, 5(2):4-8. 
4. McKinney, F.K. and Hageman, S.J. 2006. Paleozoic
to modern marine ecological shift displayed in the
northern Adriatic Sea. Geology, 34:881-884. 
3. McGowran, B., Archer, M., Bock, P., Darragh,
T.A., Godthelp, H., Hageman, S.J., Hand, S.J., Hill,
R., Li, Q., Maxwell, P.A., McNamara, K.J.,
Macaphail, M., Mildenhall, D., Partridge, A.D.,
Richardson, J., Shafik, S., Truswell, E.M., and
Warne, M. 2000. Australasian palaeobiogeography: the
Palaeogene and Neogene record. Memior of the
Association of Australasian Palaeontologists 23,
405-470. 
2. Holdener, E.J. and Hageman, S.J.
1998. Implications of intracolonial variation
in a Paleozoic bryozoan. Journal of Paleontology,
72:809-818. 
1. James, N.P., Bone, Y., Hageman, S.J., Feary, D.A.
and Gostin, V.A. 1997. Cool-water carbonate
sedimentation during the terminal Quaternary
high-amplitude, sea-level cycle: Lincoln Shelf,
southern Australia. in James, N.P. and Clark, J.A.
(eds.), SEPM Special Publication 56, Cool-water
Carbonates. 
Conference
papers and abstracts:
52. Hageman, S.J. 2009. Fossil modular
organisms (colonial invertebrates) as models
to study evolutionary rates in
developmental regulatory genes. 9th North
American Paleontological Convention Abstracts
University of Cincinnati, Cincinnati, Ohio
USA, June 21-26, 2009, p. 128.
51. Hageman, S.J. Needham, L.L. and Todd, C.D.
2008. Threshold levels for nutrient influence
on zooecia size in Electra pilosa. VIII Larwood
Meeting, Vienna, Austria, Abstracts with
Program, 17.
50. Hageman, S.J., McKinney, F.K. and Jaklin,
A. 2008. Bryozoan Colonial Growth Form and
Habitat Selection. Geological Society of
America, Annual meeting Abstracts with
Programs, 40(6): 373.
49. Steinthorsdottir, M., Wyse Jackson, P.N.,
Hageman, S.J. and Abernethy, A. 2007.
Phenotypic variation in a rhabdomesine
bryozoan from the Mississippian of
Ireland. p. 80, In: Hageman, S.J. and
McKinney, F.K (eds). Abstracts with Program,
14th Meeting of the International Bryozoology
Association, Boone, North Carolina. 2007.
48. Hageman, S.J. and Novosel, M. 2007.
Variation of zooecia size within colonies and
among environments. p. 50, In: Hageman, S.J.
and McKinney, F.K (eds). Abstracts with
Program, 14th Meeting of the International
Bryozoology Association, Boone, North
Carolina. 2007.
47. Abernethy, A. Hageman, S.J. 2007.
Environmental affects on the skeletal
phenotype of a fossil Bryozoa (Rhombopora).
Geological Society of America, Southeastern
Section, 57th annual meeting Abstracts with
Programs, 40(4): 58.
46. Steinthorsdottir, M., Wyse Jackson, P.N.,
Hageman, S.J. and Abernethy, A. 2007.
Measuring the stresses of everyday living;
Microenvironmental gradients in a rhabdomesine
bryozoan from the Mississippian of Ireland.
Abstracts with Programme, VII Larwood Meeting,
Università degli Studi di Napoli
Federico II. Naples Italy, p. 28.
45. Hageman, S.J., McKinney, F.K. 2006.
Bryozoan growth habits; complexity and
diversity in ecological reconstructions.
Geological Society of America, Southeastern
Section, 55th annual meeting Abstracts with
Programs, 38(3): 22
44. McKinney, F.K., Hageman, S.J. 2005.
Epibenthic to endobenthic transitions.
Geological Society of America, 2005 annual
meeting Abstracts with Programs, 37(7): 405.
43. Nikulina, E., Tilbrook, K.J., Sueling, J.,
Schaefer, P., Hageman, S.J. 2004. From
cryptic speciation to phylogeography in Electra pilosa;
the case of genetic study of one cosmopolitan
species. Geobiologie; 74. Jahrestagung der
Palaeontologischen Gesellschaft; Kurzfassungen
der Vortraege und Poster, 176.
42. Hageman, S.J. and Todd, C.D. 2004.
Phenotypic variation and environmental
hierarchy in the bryozoan Electra pilosa
(L.). Geological Society of America:
Abstracts with Programs v. 36.
41. Hageman, S.J., and Todd, C.D. 2004.
Hierarchical sources of environmental
variation in a modern bryozoan, Electra pilosa.
International Bryozoology Association - 13th
Conference, Concepción, 2004 –
Abstracts. Concepcion, Chile.
40. Sawyer, J.A., Hageman, S.J., and
Todd, C.D. 2003. Phenotypic variation in an
epibiotic bryozoan (Leioclema sp.) from
Chesterian ephemeral host substrates.
Geological Society of America: Abstracts
with Programs v. 35.
39. Needham, L.L., Hageman, S.J., and
Todd, C.D. 2003. Effect of food concentration
and genotype on hard-part morphology of the
marine bryozoan, Electra pilosa (L.).
Geological Society of America: Abstracts
with Programs v. 35.
38. Hageman, S.J. 2002. Paleontology and the
phenotype. Geological Society of
America: Abstracts with Programs v. 34.
37. Snyder, E.M., Hageman,
S.J., and Blake, D.B., 2002. A paleobiological
laboratory: systematics of the Carboniferous
dendroid bryozoan fauna of the Warsaw
Formation. Geological Society of
America: Abstracts with Programs v. 34.
36. Hageman, S.J., Needham, L., Bayer, M.M.,
and Todd,C.D. 2002. The influence of food
concentration on morphology (Electra pilosoa,
Bryozoa). Geological Society of America:
Abstracts with Programs v. 34, n. 7.
35. Hageman, S.J. 2002. Growth patterns and
complexity in colonial (modular) organisms.
Society for Integrated and Comparative Biology
2002 Annual Meeting and Exhibition, Final
Program with Abstracts. p. 219.
34. Hageman, S.J., Bayer, M.M., and Todd, C.D.
2001. Partitioning phenotypic variation into
Genotype, Environment and GxE Interaction and
within-colony factors: implications for
morphometric analyses (Bryozoa). Geological
Society of America: Abstracts with
Programs v. 33, n. 7.
33. Hageman, S.J., Bayer, M.M., and Todd, C.D.
2001. Partitioning sources of morphologic
variation within and among bryozoan colonies:
genetic, microenvironmental and within colony
growth/packing constraints (Electra pilosa).
12th International Bryozoology Association
Meeting, Dublin Ireland: Programme, Abstracts
& Delegates. p. 53.
32. Kazmer, M., Campbell, D., Forizs, I.,
Hageman, S.J. 2001. Bryozoan mass accumulation
on tropical latitudes: the Eocene Castle Hayne
Limestone, North Carolina, USA. 12th
International Bryozoology Association Meeting,
Dublin Ireland: Programme, Abstracts &
Delegates. p. 63.
31. Snyder, E.M., Blake, D.B., Hageman, S.J.
2001. Dendroid bryozoans of the Warsaw
Formation (Valmeyeran) of the Mississippi
River outcrop belt, Illinois, USA. 12th
International Bryozoology Association Meeting,
Dublin Ireland: Programme, Abstracts &
Delegates. p. 102.
30. Hageman, S.J. 2001. Distribution of
bryozoan growth habit characteristics in
Jacksonian (Late Eocene) facies of the Gulf
Coast. Geological Society of America,
Southeastern Section: Abstracts with
Programs v. 33, n. 2, A-15.
29. Callahan, J., Craig, J., Lusardi, W., and
Hageman, S.J. 2001. Hand specimen,
X-ray, chemical petrography, paramagnetic and
K-Ar age date studies of ballast stones from
the Queen Anne's Revenge. The Society for
Historical Archaeology, Conference on
Historical and Underwater Archaeology, Long
Beach California: Abstracts with program.
28. Hageman, S.J. and Deardorff, J.W. 2000.
Blackbeard's bryozoans and other encrusting
invertebrate fauna of ballast stones from the
Queen Anne's Revenge(?), ship wreck 03 BUI,
North Carolina. Geological Society of America,
Southeastern Section: Abstracts with
Programs v. 32.
27. Hageman, S.J., Snyder, E.M., and Holdener,
E.J. 1999. Bryozoans in morphospace
(Fenestellidae): interpreting the meaning of
fossil species. Geological Society of
America: Abstracts with Programs v. 31,
n. 7, p. 173.
26. Hageman, S.J., Lukasik, J., McGowran, B.
and Bone, Y. 1998. Paleoenvironmental signals
from Celleporaria
(Bryozoa): cyclic, cool-water carbonate
sediments from the southern Australian
Cainozoic. Geological Society of
America: Abstracts with Programs v. 31.
25. Hageman, S.J., Bock, P.E., Bone, Y. and
McGowran, B. 1998. Bryozoan growth
habits: classification and analysis.
Geological Society of America (North-central
Section): Abstracts with Programs v. 30.
24. Hageman, S.J. and Lidgard, S. 1998.
Bryozoan data exchange. 11th International
Bryozoology Association Conference,
Smithsonian Tropical Research Institute,
Panama, 1998.
23. Lidgard, S. and Hageman, S.J. 1998.
Recognizing bryozoan species. 11th
International Bryozoology Association
Conference, Smithsonian Tropical Research
Institute, Panama, 1998.
22. Beatty, W.L. Cuffey, R.J. Hageman, S.J.,
and Smith, A.M. 1998. Paleodepth and
paleohabitat from bryozoan colony forms
(Early Miocene, southern Australia).
Geological Society of America (South-eastern
Section): Abstracts with Programs v. 30,
n. 4.
21. Hageman, S.J., Bayer, M. and Todd, C.
1997. Partitioning phenotypic and genotypic
variation using the morphotype. All
Russian and International Bryozoa Conference,
St. Petersburg, Russia, 30 June-11 July, 1997.
20. James, N., Bone, Y., Hageman, S.J.,
Gostin, V.A., Feary, D.A. 1996.
The Lioness, Bonaparte's Tongue and Coo-water
Carbonates on the South Australian Continental
Margin. SEPM/AAPG Meeting.
19. Hageman, S.J., James, N., and Bone.
1996. Carbonate sediment production from
epizoic bryozoans on ephemeral
substrates. Geological Society of
America, Abstracts with Programs.
18. Hageman, S.J. 1989. Selection
of the type and number of morphometric
characters for study of fenestrate cryptostome
Bryozoa. Geological Society of
America: Abstracts with Programs v. 21,
n. 4, p. 13.
17. Hageman S.J., Bone Y., McGowran B. and
James N. 1995. Bryozoan colonial
growth forms as paleoenvironmental
indicators: reevaluation of methodology
and predictive utility. Cool and
cold-water carbonate workshop, GSA
Sedimentology Specialists Group. Melbourne
1995.
16. Hageman S.J., Bone Y., McGowran B. and
James N. 1995. Bryozoan biofacies
on the cool-water Lacepede Shelf, southern
Australia. 10th International
Bryozoology Association Conference, Wellington
New Zealand 1995.
15. Gowlett-Holmes K.L. and Hageman,
S.J., 1995. Bryozoa faunas on
jetty piles in South Australia. 10th
International Bryozoology Association
Conference, Wellington New Zealand 1995.
14. Hageman, S.J., Bone, Y., and McGowran,
B. 1994. Bryozoan facies of the
cool-water Lacepede Shelf, Southern
Australia. 12th Australian Geological
Congress, Perth, Abstracts, v. 37, p. 159.
13. Hageman, S.J. 1994.
Multivariate analysis of morphologic
variation: implications for evolutionary
studies. 12th Australian Geological
Congress, Perth, Abstracts, v. 37, p. 158.
12. Holdener, E. J., Blake, D. B., and
Hageman, S.J. 1993. Intracolonial
variation in a Mississippian fenestrate
bryozoan. Geological Society of America:
Abstracts with Programs v. 24, n. 7.
11. Holdener, E. J., Hageman, S.J., and
Blake, D. B. 1992. Intracolony
morphologic variation from a single large
fenestrate zoarium. Tenth International
Bryozoa Association Conference, Swansea Wales.
p. 23.
10. Hageman, S.J., and Blake, D. B.
1992. Microenvironmental effects on
intracolonial morphology in Bryozoa.
Geological Society of America: Abstracts with
Programs v. 24, n. 7.
9. Hageman, S.J. 1992.
Ecophenotypic and geographic constraints in a
microevolutionary study of a rhabdomesid
bryozoan. Fifth North American
Paleontological Convention, Chicago.
8. Hageman, S.J. 1992.
Implications of morphologic variability in Streblotrypa
(Rhabdomesina). Tenth International Bryozoa
Association Conference, Swansea
Wales. p. 19.
7. Hageman, S.J., Blake, D. B., and Snyder, E.
M. 1990 . From D'Arcy Thompson to
thin-plate splines: an application of
morphometrics to the systematics of Bryozoa.
Geological Society of America: Abstracts with
Programs v. 22, no 7. p. 36.
6. Hageman, S.J., and Wilkerson, M.
Scott. 1990. Non-normality and
heteroscedasticity in paleontologic data: what
is the damage done? Geological Society of
America (North-central Section):
Abstracts with Programs v. 22, n. 4, p. 202.
5. Hageman, S.J. 1989.
Evolutionary case histories: have we done the
best we can do? Geological Society of
America: Abstracts with Programs v. 22.
4. Hageman, S.J. 1989. Concepts
and methods for the taxonomic analysis of
fenestrate cryptostome Bryozoa.
International Bryozoology Association 8th
International Conference, Paris, Abstracts
Volume. Museum National d' Histoire Naturelle.
p. 14.
3. Hageman, S.J. 1989. Selection
of the type and number of morphometric
characters for study of fenestrate cryptostome
Bryozoa. Geological Society of
America: Abstracts with Programs v. 21,
n. 4, p. 13
2. Hageman, S.J. 1988. Iterative
evolution and tiering in fenestrate
Bryozoa. Geological Society of
America: Abstracts with Programs v. 20,
n. 7, p. 201
1. Hageman, S.J., and Snyder, E. M.
1987. Worthenopora
(Bryozoa, Stenolaemata): morphology and
stratigraphic usefulness of a cryptostome
(Ptilodictyina) with a cheilostome appearance.
Geological Society of America (North-central
Section): Abstracts with Programs v. 19,
n. 4, p. 202.
Unrefereed papers and reports:
5. Hageman, S.J. 2009. In Memorium: John E.
Utgaard. International Bryozoology Bulletin,
5(3): p. 8-9.
4. Hageman, S.J. 2003. Book Review: From DNA to
Diversity, Molecular Genetics and the Evolution
of Animal Design. Carroll, Grenier and
Weatherbee (2001). Journal of Paleontology.
3. Hageman S.J. 1996. Book Review:
Biology and Palaeobiology of Bryozoans, Nomen
Nudum.
2. Hageman S.J., Bone Y. and McGowran, B.
1995. Actualistic models for carbonate
production on the south Australian continental
shelf. Department of Geology and Geophysics
Annual Report 1994, The University of Adelaide,
p. 22-24.
1. Hageman S.J. 1993. Conopium
tenuissima: bryozoan epibiont from
oyster shell midden. Report for NSF funded study
on Dogan Point Shell Midden, Hudson River, New
York. Principal Investigator: Cheryl Claassen,
Dept. of Anthropology, Appalachian State
University.
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